Animals bred in zoos evolve to be less fit to survive in the wild.

This is a predictable consequence of natural selection. Animals' survival in a breeding program does not depend on the characteristics that are needed for survival in the wild, but maintaining those characteristics has costs. So they will tend to lose those characteristics.

It would be interesting to see what happens after those captivity-adapted animals are released. Do they respond to the selection pressure of wild existence by evolving in the other direction? That seems likely if enough of the original wild population remains. But if there is very little of the original wild population, the superior wild characteristics might have been lost entirely.